* {
  margin: 0;
  padding: 0;
  box-sizing: border-box;
}

body {
  background-color: #F5F4EF;
  color: #333333;
  font-family: 'PT Serif', serif;
  font-size: 14px;
  line-height: 1.7;
  padding: 40px 20px;
}

.journal-header {
  max-width: 800px;
  margin: 0 auto 30px;
}

.publication-masthead {
  display: flex;
  justify-content: space-between;
  align-items: baseline;
  margin-bottom: 8px;
}

.journal-title {
  font-family: 'Roboto Slab', serif;
  font-size: 22px;
  font-weight: 700;
  color: #556B2F;
  text-transform: uppercase;
  letter-spacing: 1px;
}

.journal-metadata {
  font-family: 'Courier Prime', monospace;
  font-size: 11px;
  color: #666;
}

.journal-divider {
  height: 3px;
  background-color: #556B2F;
  margin-bottom: 2px;
}

.journal-divider::after {
  content: '';
  display: block;
  height: 1px;
  background-color: #556B2F;
  margin-top: 2px;
}

.research-paper {
  max-width: 800px;
  margin: 0 auto;
}

.manuscript-content {
  margin-bottom: 40px;
}

.article-header {
  text-align: center;
  margin-bottom: 30px;
  padding-bottom: 20px;
  border-bottom: 1px solid #ccc;
}

.manuscript-title {
  font-family: 'Roboto Slab', serif;
  font-size: 24px;
  font-weight: 700;
  color: #556B2F;
  line-height: 1.3;
  margin-bottom: 12px;
}

.author-byline {
  font-family: 'Courier Prime', monospace;
  font-size: 11px;
  color: #666;
  font-style: italic;
}

.abstract-section {
  background-color: #EFEDE8;
  padding: 20px;
  margin-bottom: 25px;
  border-left: 4px solid #556B2F;
}

.section-label {
  font-family: 'Roboto Slab', serif;
  font-size: 14px;
  font-weight: 700;
  color: #556B2F;
  text-transform: uppercase;
  letter-spacing: 0.5px;
  margin-bottom: 10px;
}

.abstract-content {
  font-size: 13px;
  text-align: justify;
}

.navigation-module {
  margin-bottom: 25px;
  border: 1px solid #ccc;
  background-color: #FAFAF8;
}

.navigation-module details {
  padding: 15px;
}

.navigation-module summary {
  font-family: 'Roboto Slab', serif;
  font-size: 13px;
  font-weight: 700;
  color: #556B2F;
  cursor: pointer;
  text-transform: uppercase;
  letter-spacing: 0.5px;
}

.navigation-module summary:hover {
  color: #6A8EAE;
}

.navigation-module ul {
  margin-top: 12px;
  padding-left: 25px;
}

.navigation-module li {
  margin-bottom: 6px;
  font-size: 13px;
}

.navigation-module a {
  color: #6A8EAE;
  text-decoration: none;
  border-bottom: 1px solid transparent;
  transition: border-color 0.2s;
}

.navigation-module a:hover {
  border-bottom-color: #6A8EAE;
}

.methodology-block {
  column-count: 2;
  column-gap: 30px;
  text-align: justify;
}

.methodology-block .section-label {
  column-span: all;
  margin-bottom: 15px;
}

.paragraph-content {
  margin-bottom: 15px;
  break-inside: avoid;
}

.data-table-section {
  margin: 50px 0;
  break-inside: avoid;
}

.table-heading {
  font-family: 'Roboto Slab', serif;
  font-size: 14px;
  font-weight: 700;
  color: #556B2F;
  margin-bottom: 15px;
  text-align: center;
}

.comparative-data-table {
  width: 100%;
  border-collapse: collapse;
  font-size: 12px;
  background-color: #FAFAF8;
}

.comparative-data-table thead {
  background-color: #556B2F;
  color: #F5F4EF;
}

.comparative-data-table th {
  padding: 10px;
  text-align: left;
  font-family: 'Roboto Slab', serif;
  font-weight: 700;
  font-size: 11px;
  text-transform: uppercase;
  letter-spacing: 0.5px;
  border: 1px solid #556B2F;
}

.comparative-data-table td {
  padding: 10px;
  border: 1px solid #ccc;
  font-family: 'Courier Prime', monospace;
}

.comparative-data-table tbody tr:nth-child(even) {
  background-color: #F5F4EF;
}

.comparative-data-table tbody tr:hover {
  background-color: #EFEDE8;
}

.specimen-grid {
  margin: 50px 0;
}

.gallery-heading {
  font-family: 'Roboto Slab', serif;
  font-size: 14px;
  font-weight: 700;
  color: #556B2F;
  margin-bottom: 20px;
  text-align: center;
}

.photographic-evidence {
  display: grid;
  grid-template-columns: repeat(3, 1fr);
  gap: 20px;
}

.specimen-item {
  break-inside: avoid;
}

.specimen-item img {
  width: 100%;
  height: 200px;
  object-fit: cover;
  border: 2px solid #556B2F;
  display: block;
}

.figure-notation {
  font-family: 'Courier Prime', monospace;
  font-size: 11px;
  text-align: center;
  margin-top: 5px;
  color: #666;
}

.specimen-annotation {
  display: flex;
  align-items: center;
  justify-content: center;
  background-color: #EFEDE8;
  border: 2px solid #6A8EAE;
  padding: 15px;
}

.annotation-text {
  font-family: 'Courier Prime', monospace;
  font-size: 10px;
  text-align: center;
  color: #666;
  line-height: 1.5;
}

.environmental-diagram {
  margin: 50px 0;
  background-color: #FAFAF8;
  padding: 30px;
  border: 2px solid #556B2F;
}

.diagram-heading {
  font-family: 'Roboto Slab', serif;
  font-size: 14px;
  font-weight: 700;
  color: #556B2F;
  margin-bottom: 25px;
  text-align: center;
}

.schematic-flowchart {
  display: flex;
  flex-direction: column;
  align-items: center;
  gap: 0;
}

.parameter-node {
  background-color: #F5F4EF;
  border: 2px solid #6A8EAE;
  padding: 15px 25px;
  width: 100%;
  max-width: 500px;
  text-align: center;
}

.node-label {
  font-family: 'Roboto Slab', serif;
  font-size: 13px;
  font-weight: 700;
  color: #556B2F;
  margin-bottom: 5px;
}

.node-value {
  font-family: 'Courier Prime', monospace;
  font-size: 12px;
  color: #333;
}

.connector-line {
  width: 2px;
  height: 20px;
  background-color: #6A8EAE;
}

.diagram-footnote {
  font-family: 'Courier Prime', monospace;
  font-size: 10px;
  color: #666;
  margin-top: 20px;
  text-align: justify;
  font-style: italic;
}

.journal-footer {
  max-width: 800px;
  margin: 60px auto 0;
}

.footer-divider {
  height: 1px;
  background-color: #556B2F;
  margin-bottom: 15px;
}

.footer-content {
  display: flex;
  justify-content: space-between;
  align-items: center;
  font-family: 'Courier Prime', monospace;
  font-size: 11px;
  color: #666;
}

.contact-information a {
  color: #6A8EAE;
  text-decoration: none;
}

.contact-information a:hover {
  text-decoration: underline;
}

.page-notation {
  font-style: italic;
}

@media (max-width: 768px) {
  body {
    padding: 20px 15px;
  }

  .journal-title {
    font-size: 18px;
  }

  .manuscript-title {
    font-size: 20px;
  }

  .methodology-block {
    column-count: 1;
  }

  .photographic-evidence {
    grid-template-columns: repeat(2, 1fr);
    gap: 15px;
  }

  .specimen-item img {
    height: 150px;
  }

  .footer-content {
    flex-direction: column;
    gap: 10px;
    text-align: center;
  }

  .parameter-node {
    padding: 12px 15px;
  }

  .node-label {
    font-size: 12px;
  }

  .node-value {
    font-size: 11px;
  }
}

@media (max-width: 480px) {
  .photographic-evidence {
    grid-template-columns: 1fr;
  }
}